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What is a custos?

What is a custos? Here are some definitions.

Noun
  1. (obsolete) A warden.
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Examples
Should he go direct to Mount Welcome and warn the custos, by reporting to him what he had heard?
Melody-Harmony generates and displays custos automatically.
Edward crossed the English Channel to France in January, leaving Gaveston as his custos regni in charge of the kingdom in his absence.
Usually, though not necessarily, the person appointed lieutenant was also appointed custos rotulorum or keeper of the rolls.
The custos read it all over slowly and carefully, weighing every word.
The custos rotulorum also appointed the clerk of the peace, who advised the justices on legal matters beyond the scope of the quorum.

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